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Rubus hawaiensis

A.Gray (1854)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Rosaceae Genus: Rubus

‘ākala [akala], ‘ākalakala [akalakala], kala

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Description

Key Characters:

Growth Form: Shrubs.

Stems: Shredding bark, stems erect or arching, up to 40 dm long, sparingly branched, primocanes and floricanes similar, covered with slender prickles up to 4 mm long, the bases sometimes elongated longitudinally, or nearly as often without prickles, also sparsely to moderately or sometimes densely pilose.

Roots:

Leaves: Leaves compound (palmate). Alternate. Blades ovate to rhombic–ovate or deltate–ovate, the lateral ones slightly asymmetrical, leaflets 3, the terminal one largest, 8–15(–21) cm long, 5–10(–15) cm wide., thin, petiolules (1.3–)2.5–3.5 cm long, petiolules sparsely pilose to densely tomentose, sometimes covered with short slender prickles. Upper surfaces pilose to tomentose along the impressed veins; lower surfaces densely or sometimes moderately tomentose, sometimes pale, sparsely short-pilose only along veins. Margins coarsely and irregularly serrate and also often somewhat lobed. Petioles sparsely pilose to densely tomentose, sometimes covered with short slender prickles. Stipules adnate to base of petioles, linear–lanceolate, 1–1.8 cm long.

Flowers: Flowers 1–4(–8) in corymbose inflorescences. Flowers bisexual (perfect), actinomorphic; pedicels 13–30 mm long, pilose to tomentose. Receptacle raised, margins bearing sepals, petals, and stamens. Calyx lobes 5, persistent, sepals ovate, distinct, imbricate, 8–17 mm long, both surfaces sparsely to densely tomentose, apex long–acuminate or caudate. Corolla of 5 petals, dark pink to dusky rose or rarely white, obovate, distinct, imbricate, ca. 13–18 mm long. Stamens numerous; filaments slender; anthers dithecal. Ovary superior, inserted on the raised receptacle, 1-celled; ovules 2 per cell; style filiform, subterminal, persistent consisting of coherent.

Fruit: Fruit fleshy; juicy drupelets; red to dark purple or occasionally yellow; ovoid; ca. 2.5–5 cm long. Seeds 1 per drupelet.

Ploidy:

Habitat: Mesic to wet forest and subalpine woodland.

Elevation Range: 660–3070 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Only found in cultivation
Molokai Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: U.S. Expl. Exped., Phan. 1: 504 (1854)

Other References

Wagner et al. 1990:1109 (K, Mo, M, H); Note: this was planted at the Ka‘ala summit on O at one point but there are no known natural occurrences on the island

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Rubus hawaiensis Kulani open, shrubby forest Preserved_Specimen no prickles Degener, O. 30821 Hawaii BISH 1966-05-26
2 Rubus hawaiensis Pihea trail wet forest Preserved_Specimen ~2 m shrub, sprawling; corolla pink Wagner, W.L. 6034 Kauai BISH 1988-04-09
3 Rubus hawaiensis Hawaii National Park, Kipuka Kulalio SE exposure in Koa parklands, entrance to lava tube where protected from cattle grazing and trampling Preserved_Specimen 4'; flower pink, berry yellow Baldwin, P.H. 78 Hawaii BISH 1941-09-17
4 Rubus hawaiensis Puu o Kaeha, Kawela wooded stream Preserved_Specimen 10' long canes, arching St.John, H. 12464 Molokai BISH 1932-12-23
5 Rubus hawaiensis Maui in the Deschampsia-Holcus ecotone below Vogl bog, Haleakala. Preserved_Specimen Takeuchi, W.N. 5 Maui BISH 1983-08-24
6 Rubus hawaiensis Keanakolu, Humuula, Hilo FR moist gulch, Acacia koa forest Preserved_Specimen 2-3 m shrub, flower pink St.John, H. 18597 Hawaii BISH 1937-12-30
7 Rubus hawaiensis Laupahoehoe section of Hilo Forest Reserve, betw E fork of Kaawali stream & NW of Haakoa stream Open ohia forest with Myrsine, Gouldia, Clermontia parviflora, Cheirodendron, Ilex. Preserved_Specimen Shrub 5ft tall, flowers pink. Davis, J. 692 Hawaii BISH 1982-02-19
8 Rubus hawaiensis N Hilo, 38.4 km along Saddle Rd. under Metrosideros, shady Preserved_Specimen large shrub Herat, T. 745 Hawaii BISH 1974-06-22
9 Rubus hawaiensis Laumaia koa, lehua forest Preserved_Specimen Neal, M.C. 805 Hawaii BISH 1935-08-17
10 Rubus hawaiensis East Maui, Koolau Forest Reserve In tussock grassland near Puu Alaea. In small gulch with Coprosma, Sophora, and Vaccinium Preserved_Specimen Harrison, B. 496 Maui BISH 1973-10-10
11 Rubus hawaiensis Maui in the Deschampsia-Holcus ecotone below Vogl bog, Haleakala. Preserved_Specimen Takeuchi, W.N. 5 Maui BISH 1983-08-24
12 Rubus hawaiensis Olinda, above grassy slopes by stream bank Preserved_Specimen shrub 8 ft. tall St.John, H. 10335 Maui BISH 1930-02-13
13 Rubus hawaiensis Haleakala National Park, Palikou [= Paliku], behind cabin grasslands Preserved_Specimen Large bush up to 3 m., just beginning to lfower and leaf. Stems white to light brown. Spines small, light brown. Leaves light green. Calyx light green. Flowers pale purple, fragrant. Fruits immature, dark purple-red. van Royen, P. 11854 Maui BISH 1981-03-23
14 Rubus hawaiensis Makawao District, Haiku - Uka Trail, Haleakala Wet forest Preserved_Specimen Canes ascending 2 meters tall. Fosberg, F.R. 9855 Maui BISH 1933-08-18
15 Rubus hawaiensis Keanakolu, Humuula, Hilo FR moist gulch, Acacia koa forest Preserved_Specimen 2-3 m shrub, flowers white St.John, H. 18598 Hawaii BISH 1937-12-30
16 Rubus hawaiensis Kokee, Waimea, Napali-Kona FR wet woods Preserved_Specimen St.John, H. 10691 Kauai BISH 1930-12-26
17 Rubus hawaiensis Hilo, Puu Oo trail above Olaa flume kipuka in 1881 flow Preserved_Specimen 3 m arching cane; sterile Fosberg, F.R. 10450 Hawaii BISH 1933-12-10
18 Rubus hawaiensis Puu Huluhulu, Humuula, N Hilo hill kipuka, open forest Preserved_Specimen 2 m arching shrub St.John, H. 23933 Hawaii BISH 1949-12-24
19 Rubus hawaiensis Keauhou Ranch, Kau District. (T-KU) Log jam with Metrosideros and Cibotium surrounded by Pennisetum - Axonopus - Anthoxanthum pasture. Preserved_Specimen Height: 8 ft. Flowers purplish pink. Cuddihy, L.W. 151 Hawaii BISH 1980-01-29
20 Rubus hawaiensis Hawaii National Park, Kipuka Kulalio SE exposure in Koa parklands, entrance to lava tube where protected from cattle grazing and trampling Preserved_Specimen 4'; flower pink, berry yellow Baldwin, P.H. 78 Hawaii BISH 1941-09-17
21 Rubus hawaiensis Piihonua; Hilo Forest Reserve, Site 22 OS Open healthy forest; assoc. spp.: Metrosideros polymorpha, Ilex anomala, Gouldia sp., Pelea clusiifolia, Cheirodendron trigynum, Myrsine lessertiana, Freycinetia arborea, Alyxia oliviformis. Soil a deep moderately well drained ash. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-19
22 Rubus hawaiensis Puu Makaala Natural Area Reserve Site Along unpaved road on edge between volcanic substrates of different ages in Acacia koa forest. Preserved_Specimen Fruiting Preer, S. 14 Hawaii BISH 2020-07-13
23 Rubus hawaiensis Kula Pipe Line, Olinda thicket by trail Preserved_Specimen St.John, H. 10293 Maui BISH 1930-02-11
24 Rubus hawaiensis NW slopes of Haleakala, Kula Forest Reserve, lower part moist shrubby gullies Preserved_Specimen shrub to 3.5 m, flowers pink, ripe fruit red, stamens persistent, red in fruiting stage; frequent. Stauffer, H.U. 5886 Maui BISH 1964-04-17
25 Rubus hawaiensis Olokui, Wailele Valley rain forest Preserved_Specimen 5 m tall canes, straight St.John, H. 23252 Molokai BISH 1948-02-04
26 Rubus hawaiensis Keauhou Ranch, Kau District, Puu Kipu In closed Metrosideros - Acacia koa forest. Preserved_Specimen Very common understory shrub. Height 10 ft. Flowers purplish pink. Cuddihy, L.W. 196 Hawaii BISH 1980-02-12
27 Rubus hawaiensis E end Haleakala Crater behind Paliku Ranger Cabin Preserved_Specimen Forming thicket with shrubs to 7 feet tall, petals lavender, fruit dark purple-red when ripe, filaments red. Henrickson, J.S. 3716 Maui BISH 1969-06-23
28 Rubus hawaiensis 6.75 mi E of Puu Huluhulu, Saddle Rd. moss, Metrosideros forest, kipuka Preserved_Specimen St.John, H. 25342 Hawaii BISH 1954-12-25
29 Rubus hawaiensis S Kona, McCandless Ranch rain forest Preserved_Specimen stems prickly Rock, J.F.C. 27511 Hawaii BISH 1959-03-22
30 Rubus hawaiensis Keahou Ranch, Puu Oo, S Mauna Kea small wooded ravine, pasture, near house Preserved_Specimen Fosberg, F.R. 59510 Hawaii BISH 1980-03-08
31 Rubus hawaiensis Mauna Loa, above Kulani rain forest Preserved_Specimen Christ, J.H. 27 Hawaii BISH 1957-04-24
32 Rubus hawaiensis Drive to end of Kulani Prison Road about 3 miles past Kulani Prison (Honor) Camp; or 19 miles from intersection with Kona-Hilo Highway near Hilo Mixed Acacia - Metrosideros - Cibotium community Preserved_Specimen Shrub about 1 foot tall, creeper Newell, C.L. 499 Hawaii BISH 1966-08-21
33 Rubus hawaiensis Haleakala National Park, Palikou [= Paliku], behind cabin grasslands Preserved_Specimen Large bush up to 3 m., just beginning to lfower and leaf. Stems white to light brown. Spines small, light brown. Leaves light green. Calyx light green. Flowers pale purple, fragrant. Fruits immature, dark purple-red. van Royen, P. 11854 Maui BISH 1981-03-23
34 Rubus hawaiensis Near hill northeast of Olinda Sheltered open thicket in catle range. Preserved_Specimen Degener, O. 2259 Maui BISH 1927-06-15
35 Rubus hawaiensis East Maui, Hana District, Waihoi Valley South Fork of Waiohonu at base of a 250 ft waterfall. With Dubautia sp. and Perrottetia. Preserved_Specimen Harrison, B. 35 Maui BISH 1972-06-14
36 Rubus hawaiensis East Maui, Hana District, Keanae AHU, Tr 4(43), N slope of Haleakala, Keanae Valley Wet 'ohi'a forest with other native trees and a treefern and native shrub understory. Preserved_Specimen Leaves w/concentric rings Warshauer, F.R. 2825 Maui BISH 1980-07-14
37 Rubus hawaiensis Laumaia koa, lehua forest Preserved_Specimen Neal, M.C. 805 Hawaii BISH 1935-08-17
38 Rubus hawaiensis Mauna Kea, along Wailuku River road near State silversword exclosure Transitional Sophora forest Styphelia scrub. Preserved_Specimen Excellent vigor. Shrub 12' height, 10' spread, 1 basal diameter. Flowering profusely. Clarke, G. 434 Hawaii BISH 1980-06-03
39 Rubus hawaiensis Puu Huluhulu, Humuula Sheep Station Acacia koa, Coprosma, Myoporum; shade Preserved_Specimen erect shrub with lax, arching branches to 9' long; foliage dull, dark green, lighter below; flower deep rose; fruit red; bark at base orangish brown, dry, thin, papery, scaly Herbst, D.R. 914 Hawaii BISH 1968-02-13
40 Rubus hawaiensis Wright Road, Olaa Forest Reserve, Puna Cibotium and Metrosideros forest Preserved_Specimen 7-10' arching cane; petals deep pink St.John, H. 25081 Hawaii BISH 1953-01-29
41 Rubus hawaiensis Puu Huluhulu, Humuula Sheep Station Acacia koa, Coprosma, Myoporum; shade Preserved_Specimen erect shrub with lax, arching branches to 9' long; foliage dull, dark green, lighter below; flower deep rose; fruit red; bark at base orangish brown, dry, thin, papery, scaly Herbst, D.R. 914 Hawaii BISH 1968-02-13
42 Rubus hawaiensis 4 mi due E of Olinda on Haipuaena Gulch Deep canyon-like gulch at head of Olinda water flume; stream sides and rocky cliffs and sterep lush slopes, with Gunnera mauiensis, Cyanea ferox, Rubus hawaiensis, Broussaisia arguta. Preserved_Specimen 2m. tall, none in flower or fruit now. No thorns. Rocky streambed. Bonsey, E. 454 Maui BISH 1967-08-05
43 Rubus hawaiensis Saddle Rd., 29 miles, 1855 lava flow, Waiakea, S Hilo wooded kipuka Preserved_Specimen 6 m cane, arching St.John, H. 22392 Hawaii BISH 1946-12-26
44 Rubus hawaiensis East Maui, Ukulele In the forest. Preserved_Specimen Forbes, C.N. 957 Maui BISH 1919-07-01
45 Rubus hawaiensis Waimea District central Alakai, headwaters of Koaie stream, northwestern portion of Drinking Glass proposed fence unit Metrosideros-Cheirodendron montane wet forest with Dicranopteris- Diplopterygium-Sticherus matting ferns with Leptecophylla tameiameiae, Vaccinium calycinum, Broussaisia arguta, Kadua affinis, Peperomia hesperomannii, Gahnia vitiensis kauaiensis, Coprosma elliptica, Melicope cluusiifolia, M. Preserved_Specimen shrub, diffusely branched, with red-purple fruit, occasional Wood, K.R. 15516 Kauai BISH 2013-05-22
46 Rubus hawaiensis Waikolu Valley, head shrubby, cloud Preserved_Specimen Degener, O. 22170 Molokai BISH 1952-03-15
47 Rubus hawaiensis Olaa Flume Preserved_Specimen Forbes, C.N. 645 Hawaii BISH 1915-06-02
48 Rubus hawaiensis Kawaikoi Stream, Mohihi road, forestry cabin Preserved_Specimen Takeuchi, W.N. 96 Kauai BISH 1983-04-11
49 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
50 Rubus hawaiensis Kaala, Waianae Mts Preserved_Specimen 3 m; 3 flowers St.John, H. 22271 Oahu BISH 1946-11-17
51 Rubus hawaiensis Kipuka Ahiu Preserved_Specimen Forbes, C.N. 897 Hawaii BISH 1915-06-21
52 Rubus hawaiensis East Maui, Kaupo Gapl Crater of Haleakala Preserved_Specimen Forbes, C.N. 1117 Maui BISH 1919-08-10
53 Rubus hawaiensis Kau District, Kilauea Forest, along TKH Preserved_Specimen 5 feet tall. Kubo, R. 56 Hawaii BISH 1980-02-26
54 Rubus hawaiensis Kamoku Preserved_Specimen Forbes, C.N. 686 Molokai BISH 1915-03-31
55 Rubus hawaiensis Kipahulu Valley, central pali of valley Preserved_Specimen Plants 2 m tall, growing on steep slopes, forming clumps 3-4 m in diameter. Lamoureux, C.H. 3924 Maui BISH 1967-08-07
56 Rubus hawaiensis Poholua Preserved_Specimen Forbes, C.N. 89 Molokai BISH 1912-06-01
57 Rubus hawaiensis West slope of Haleakala Preserved_Specimen Bright pink flowers and enormous fruits, the drupes ca. 7mm in diameter. The entire fruit 1 1/2 X 1 1/4 at time picked. Stamens formimng a corona at base of fruit. Stamen filaments turn pink with the ripening of the fruits. Fruits mildly sour with no fragrance. Gillett, G.W. 1559 Maui BISH 1964-04-17
58 Rubus hawaiensis Kalalau Trail Preserved_Specimen Forbes, C.N. 1086 Kauai BISH 1917-07-03
59 Rubus hawaiensis National Park, trail from Mauna Loa scenic road shelter Preserved_Specimen aggregate fruit Herat, T. 635 Hawaii BISH 1974-06-19
60 Rubus hawaiensis East Maui, Makawao Preserved_Specimen Munro, G.C. 485 Maui BISH 1918-04-24
61 Rubus hawaiensis Puu Huluhulu Preserved_Specimen fruit red Forbes, C.N. 832 Hawaii BISH 1915-06-12
62 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 914 Maui BISH 1919-07-01
63 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
64 Rubus hawaiensis Kalaupapa; Puu Alii, Pelekunu rim below trail, 100 m S of park fence Preserved_Specimen shrub Wysong, M.L. 656 Molokai BISH 2005-03-22
65 Rubus hawaiensis above Humuula, Hookomo erosion crack Preserved_Specimen Neal, M.C. 741 Hawaii BISH 1935-08-08
66 Rubus hawaiensis Waikolu Valley, head and E rim Preserved_Specimen ~7' shrub, no spines Kondo, Y. Molokai BISH 1962-06-18
67 Rubus hawaiensis Waikolu Valley, head and E rim Preserved_Specimen ~7' shrub, no spines Kondo, Y. Molokai BISH 1962-06-18
68 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
69 Rubus hawaiensis Gulch near Hosmer Nursery Preserved_Specimen Krajina, V.J. 620415042 Maui BISH 1962-04-15
70 Rubus hawaiensis Kaholuamanu Preserved_Specimen Rock, J.F.C. 2202 Kauai BISH 1909-03-03
71 Rubus hawaiensis Glenwood Preserved_Specimen Faurie, A. 802 Hawaii BISH 1909-05-01
72 Rubus hawaiensis Kaala, Waianaeuka Preserved_Specimen 4 m shrub St.John, H. 20130 Oahu BISH 1941-01-12
73 Rubus hawaiensis National Park, trail from Mauna Loa scenic road shelter Preserved_Specimen aggregate fruit Herat, T. 635 Hawaii BISH 1974-06-19
74 Rubus hawaiensis East Maui, Hana District, E end Haleakala Crater, behind Paliku Ranger Cabin Preserved_Specimen Forming thicket with shrubs to 7 feet tall, petals lavendar, fruit dark purple-red when ripe, filaments red. Henrickson, J.S. 3716 Maui BISH 1969-06-23
75 Rubus hawaiensis Puu Huluhulu Preserved_Specimen fruit yellow Forbes, C.N. 833 Hawaii BISH 1915-06-12
76 Rubus hawaiensis Kamoku-Waikolu Valley Preserved_Specimen Rock, J.F.C. 6189 Molokai BISH 1910-03-28
77 Rubus hawaiensis Kalahuapueo to Hanalilolilo Preserved_Specimen Krajina, V.J. 620611101 Molokai BISH 1962-06-11
78 Rubus hawaiensis Waikolu Valley, head and E rim Preserved_Specimen ~7' shrub, no spines Kondo, Y. Molokai BISH 1962-06-18
79 Rubus hawaiensis trail between Halelouolu and Wailuku River Preserved_Specimen Forbes, C.N. 715 Hawaii BISH 1915-06-05
80 Rubus hawaiensis Kawaikoi Stream, Mohihi road, forestry cabin Preserved_Specimen Takeuchi, W.N. 96 Kauai BISH 1983-04-11
81 Rubus hawaiensis [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Faurie, A. 1 BISH 1909-01-01
82 Rubus hawaiensis East Maui, Hamakua Preserved_Specimen Hillebrand, W.B. Maui BISH
83 Rubus hawaiensis E of Ukulele Preserved_Specimen Forbes, C.N. 727 Maui BISH 1919-07-11
84 Rubus hawaiensis Paauhau Preserved_Specimen Rock, J.F.C. 8305 Hawaii BISH 1910-05-01
85 Rubus hawaiensis E of Ukulele Preserved_Specimen Forbes, C.N. 727 Maui BISH 1919-07-11
86 Rubus hawaiensis Paauhau Preserved_Specimen Rock, J.F.C. 8305 Hawaii BISH 1910-05-01
87 Rubus hawaiensis Kokee Camp Preserved_Specimen Degener, O. 30190 Kauai BISH 1976-06-23
88 Rubus hawaiensis Waihualele Gulch, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1827 Maui BISH 1920-03-04
89 Rubus hawaiensis 17 mi from Kona toward Waimea, main road Preserved_Specimen Degener, O. 30189 Hawaii BISH 1976-08-14
90 Rubus hawaiensis Puuouli, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2152 Maui BISH 1920-04-05
91 Rubus hawaiensis Mauna Kea, Waikii Preserved_Specimen Rock, J.F.C. 8331 Hawaii BISH 1910-06-01
92 Rubus hawaiensis Kamoku Preserved_Specimen Rock, J.F.C. 16014 Molokai BISH 1918-04-01
93 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 914 Maui BISH 1919-07-01
94 Rubus hawaiensis Glenwood Preserved_Specimen Faurie, A. 802 Hawaii BISH 1909-05-01
95 Rubus hawaiensis Kamalo Preserved_Specimen Faurie, A. 805 Molokai BISH 1910-07-01
96 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
97 Rubus hawaiensis Glenwood Preserved_Specimen Faurie, A. 802 Hawaii BISH 1909-05-01
98 Rubus hawaiensis Honauma [Maunahooma?, W. Maui Preserved_Specimen Forbes, C.N. 9 Maui BISH 1910-05-01
99 Rubus hawaiensis Kipuka Puauluu, Kilauea Preserved_Specimen Rock, J.F.C. 12567 Hawaii BISH 1915-07-01
100 Rubus hawaiensis Kamalo Preserved_Specimen Faurie, A. 805 Molokai BISH 1909-06-01
101 Rubus hawaiensis S Kona, Koa Camp Preserved_Specimen Skottsberg, C.J.F. 1927 Hawaii BISH 1926-09-23
102 Rubus hawaiensis East Maui, woods near Ukulele, above Olinda Preserved_Specimen Forbes, C.N. 168 Maui BISH 1910-07-01
103 Rubus hawaiensis Wailau Valley Preserved_Specimen Forbes, C.N. 560 Molokai BISH 1912-09-01
104 Rubus hawaiensis Kaholuamanu Preserved_Specimen Rock, J.F.C. 2201 Kauai BISH 1909-03-03
105 Rubus hawaiensis Waihualele Gulch, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1827 Maui BISH 1920-03-04
106 Rubus hawaiensis East Maui, Makawao Preserved_Specimen Munro, G.C. 666 Maui BISH 1918-04-24
107 Rubus hawaiensis Mauna Kea, Waikii Preserved_Specimen Rock, J.F.C. 8331 Hawaii BISH 1910-06-01
108 Rubus hawaiensis Kamoku Preserved_Specimen Forbes, C.N. 686 Molokai BISH 1915-03-31
109 Rubus hawaiensis Mauna Kea, Waikii Preserved_Specimen Rock, J.F.C. 8331 Hawaii BISH 1910-06-01
110 Rubus hawaiensis Above Waikolu Preserved_Specimen Munro, G.C. 386 Molokai BISH 1916-06-05
111 Rubus hawaiensis Laumaia Preserved_Specimen Forbes, C.N. 890 Hawaii BISH 1915-06-18
112 Rubus hawaiensis Haleakala Preserved_Specimen Faurie, A. 806 Maui BISH 1909-08-01
113 Rubus hawaiensis Back of Kilauea Preserved_Specimen Lydgate, J.M. Hawaii BISH 1909-07-01
114 Rubus hawaiensis 200-300 yds, E of Kalalau Valley Lookout, Waimea, Kokee Preserved_Specimen Newhouse, W.J. Kauai BISH 1959-05-03
115 Rubus hawaiensis East Maui, Hamakua Preserved_Specimen Hillebrand, W.B. Maui BISH
116 Rubus hawaiensis Waiakoali Preserved_Specimen Neal, M.C. Kauai BISH 1930-08-22
117 Rubus hawaiensis National Park, Puu Huluhulu Crater Preserved_Specimen Kemp, R.M. Hawaii BISH 1975-08-06
118 Rubus hawaiensis Kauai Preserved_Specimen Seale, A. Kauai BISH
119 Rubus hawaiensis upper Maunahui camp to Pepeopae bog Preserved_Specimen Cranwell, L.M. 2506 Molokai BISH 1938-07-12
120 Rubus hawaiensis Puuouli, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2152 Maui BISH 1920-04-05
121 Rubus hawaiensis Kipahulu Valley, Haleakala National Park, National Park Service Expedition III, West Camp, transect 5 Preserved_Specimen Russell, C.A. 561 Maui BISH 1984-03-03
122 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
123 Rubus hawaiensis Above Waikolu Preserved_Specimen Munro, G.C. 573 Molokai BISH 1916-06-05
124 Rubus hawaiensis S Kula slope of Haleakala Preserved_Specimen Forbes, C.N. 2170 Maui BISH 1920-04-07
125 Rubus hawaiensis Ukulele Preserved_Specimen Forbes, C.N. 972 Maui BISH 1919-07-29
126 Rubus hawaiensis E of Ukulele Preserved_Specimen Forbes, C.N. 727 Maui BISH 1919-07-11
127 Rubus hawaiensis unknown Preserved_Specimen Faurie, A. 1 BISH 1909-01-01
128 Rubus hawaiensis [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Faurie, A. 1 BISH 1899-01-01
129 Rubus hawaiensis [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Hillebrand, W.B. BISH
130 Rubus hawaiensis Kona Preserved_Specimen Mann, H. 576 Hawaii BISH 1864-01-01
131 Rubus hawaiensis Honauma [Maunahooma?, W. Maui Preserved_Specimen Forbes, C.N. 91 Maui BISH 1910-05-01
132 Rubus hawaiensis West Ridge of Haipuaena Preserved_Specimen Forbes, C.N. 2632 Maui BISH 1920-06-28
133 Rubus hawaiensis Papaloa, Kona Preserved_Specimen Forbes, C.N. 309 Hawaii BISH 1911-07-14
134 Rubus hawaiensis trail between Halelouolu and Wailuku River Preserved_Specimen Forbes, C.N. 715 Hawaii BISH 1915-06-05
135 Rubus hawaiensis David Douglas Monument shaded, rock outcrops, koa forest Preserved_Specimen 4-7' erect stems; fruit orange to red Tomich, P.Q. 932 Hawaii BISH 1962-03-27
136 Rubus hawaiensis Waikolu Valley, head shrubby, cloud Preserved_Specimen Degener, O. 22170 Molokai BISH 1952-03-15
137 Rubus hawaiensis Puu Kukui Rain forest Preserved_Specimen Shrub 1.5 meters tall, petals pink, fruit dark red at maturing. Wilbur, R.L. 870 Maui BISH 1948-06-28
138 Rubus hawaiensis Makawao District, Ahupuaa of Kalialinui, upper Koolau Gap Open Styphelia-Vaccinium dominated pahoehoe W of Waikau Cabin. Big patch at base of cliff behind Waikau Cabin. Preserved_Specimen Berries purple and red. Higashino, P.K. 772 Maui BISH 1975-06-01
139 Rubus hawaiensis Makawao District, Haiku - Uka Trail, Haleakala Wet forest Preserved_Specimen Flowers pink, long drooping canes. Fosberg, F.R. 9877 Maui BISH 1933-08-18
140 Rubus hawaiensis Mauka of Kulani Prison open forest Preserved_Specimen Degener, O. 21824 Hawaii BISH 1952-02-15
141 Rubus hawaiensis Kokee, Waimea, Napali-Kona FR wet woods Preserved_Specimen St.John, H. 10691 Kauai BISH 1930-12-26
142 Rubus hawaiensis above Halealoha, 1855 flow kipuka Preserved_Specimen Forbes, C.N. 786 Hawaii BISH 1915-06-09
143 Rubus hawaiensis Hilo, Puu Oo trail above Olaa flume kipuka in 1881 flow Preserved_Specimen 3 m arching cane; sterile Fosberg, F.R. 10450 Hawaii BISH 1933-12-10
144 Rubus hawaiensis Wai Anapanapa, Haleakala, Kipahulu-Kuhiwa divide thicket just above lake flat Preserved_Specimen common canes mostly upright, 2-3 m. high. leaves membranous, bright green, fruit 1.5-2.5 cm. long, dark cherry red, shiny, glabrous except for sparse capitate puberulence near tip. St.John, H. 20992 Maui BISH 1945-08-14
145 Rubus hawaiensis Keauhou Ranch, Kau District, T-KD. Koa reforestation area, in bulldozed increment 3 In bulldozed area. Preserved_Specimen Height 4 ft. No flowers, fruit red- purple. Cuddihy, L.W. 123 Hawaii BISH 1980-01-14
146 Rubus hawaiensis Puu Huluhulu, Humuula Sheep Station Acacia koa, Coprosma, Myoporum; shade Preserved_Specimen erect shrub with lax, arching branches to 9' long; foliage dull, dark green, lighter below; flower deep rose; fruit red; bark at base orangish brown, dry, thin, papery, scaly Herbst, D.R. 914 Hawaii BISH 1968-02-13
147 Rubus hawaiensis Ainahou aa clinker, ohia Preserved_Specimen arching stems Takeuchi, W.N. 3655 Hawaii BISH 1987-08-01
148 Rubus hawaiensis Maui Disturbed, predominantly open scrub, (Sophora, Styphelia)-dominated, slope S. of Hosmer's Grove Preserved_Specimen Higashino, P.K. 661 Maui BISH 1975-06-12
149 Rubus hawaiensis Kawaiiki Valley, below Kaluahaulu ridge trail, to th e W of the Diellia fence, in a side gulch with Labordia forencia Mixed Mesic Forest canopy of Acacia koa and Metrosideros polymorpha w/Dicranopteris, Labordia lorenciana, Scaevola, Tetraplasandra, Cheirodendron, Wikstroemia furcatra, Alyxia stellata, Elaeocarpus bifidus, Dianella sandwicensis, Bidens cosmoides, Dubautia, Kadua affinis, Psychotria, Dryopteris. Preserved_Specimen Large shrub covering 10 x 10 patch. 6 ft tall sprawling. Fls white, petals pink with age and falling. Lvs light green-yuellow. Bark red-brown with tan lifting segments. Tangalin, N. 1970 Kauai BISH 2009-04-08
150 Rubus hawaiensis National Park, Kipuka Puaulu dry forest Preserved_Specimen 2 m viney shrub; petals pink Webster, G.L. 1789 Hawaii BISH 1948-06-18
151 Rubus hawaiensis East Maui, Waikamoi rain forest Preserved_Specimen Neal, M.C. Maui BISH 1933-08-14
152 Rubus hawaiensis N Hilo, Saddle Rd, Kipuka Ainahou, 30 mi SSE of jct with Hwy. 190 with Metrosideros, Vaccinium, Styphelia, Coprosma Preserved_Specimen 1.5 m shrub, erect arcuate stems; petals pale purplish pink; ripe fruit red, juicy Lorence, D.H. 5521 Hawaii BISH 1987-08-07
153 Rubus hawaiensis Mauna Loa strip road open scrub Preserved_Specimen 2 m many stemmed shrub, canes arching; flower rose pink; ripe fruit globose, scarlet, 2-2.5 cm diam. Fosberg, F.R. 57484 Hawaii BISH 1978-04-14
154 Rubus hawaiensis Puu Hualalai, Puuwaawaa moist woods Preserved_Specimen St.John, H. 11360 Hawaii BISH 1931-12-29
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